A new species Theronia qaidamensis sp. nov. (Ichneumonidae, Pimplinae) is described from the Oligocene Shangganchaigou Formation of the northwestern Qaidam Basin (Huatugou area), northeast of the Tibetan Plateau (TP). The new species represents the first report of the Ichneumonidae from the Qaidam Basin, which reveals an insect linkage between the Qaidam Basin and Europe. In addition, the study of associated fossils from the Shangganchaigou Formation shows that the palaeoenvironment of the northwestern Qaidam Basin during the Oligocene was warm and moist, which provided suitable living conditions for the current fossil.
Fossils are critical for understanding the environmental changes and uplift history of the Tibetan Plateau during the Cenozoic. As the largest inland basin located at the northeastern plateau, Qaidam Basin has been regarded as the key region to understand the climatic changes in the plateau during the Cenozoic. Herein, fossil legumes from the Oligocene Shangganchaigou Formation in northwestern Qaidam Basin have been reported and are assigned to Desmanthus cf. virgatus (L.) Willd. based on morphological comparison. Current materials represent the first megafossil record of this genus in the world. The living analogues of current fossils now live in regions with mean annual temperature values (MAT) ranging from 13.4 to 29.1°C and mean annual precipitation values (MAP) ranging from 36 to 2874 mm while the MAT and MAP of current northwestern Qaidam Basin is −1.03°C and 60.5 mm, respectively, and indicate that the Oligocene Qaidam was much warmer than today. We calculate that current fossils could grow no higher than 2492 m considering the climatic condition of the Oligocene Qaidam. The difference between estimated and modern elevation of the fossil site suggests that the elevation of the research area has increased by at least ≈1000 m since the Oligocene.
Trapa L. (Lythraceae J. ST.-HIL) is a genus of annual free-floating plant, widely distributed in the Northern Hemisphere. The Qaidam Basin is regarded as an ideal region to understand the Cenozoic palaeoclimatic and palaeobiological evolution of the northern Tibetan Plateau. A few records of Trapa have been previously mentioned from the Cenozoic deposits of the Qaidam Basin, northern Tibetan Plateau. Here, we describe some fossil fruits of Trapa from the late Miocene Shangyoushashan Formation of southeastern Qaidam Basin (Tuosu Lake area), which were identified as Trapa qaidamensis sp. nov. based on their morphology. Extant species of Trapa usually live in the environments with mean temperature of warmest quarter (MTWQ) ranging from 14.9°C to 30.9°C, as well as mean temperature of warmest month (MTWM) ranging from 19.6°C to 37.3°C in the living areas of Trapa. Compared with MTWQ 16.5°C and MTWM 17.5°C in current fossil locality, the present fossil site was warmer during the Miocene than in the present day.urn:lsid:zoobank.org:act:9F7DAD1B-DA2D-4734-BE27-99276F57DD2A
Equisetum (Equisetaceae, Equisetales) is widely distributed around the world and distinguished by obviously jointed stems with longitudinal ridges or furrows. Recently, fossil materials collected from the Shangganchaigou Formation of northwestern Qaidam Basin, China, are assigned to Equisetum based on the presence of jointed stems, longitudinal ridges, and whorls of branch scars. The current specimens are the first discovery of Equisetum in northern Tibetan Plateau and also represent the first fossil record of the genus from Oligocene in Qaidam Basin. The new discovery enriches the knowledge of the paleo-diversity of the Equisetum and indicates that the sphenophytes once existed in the northern Tibetan Plateau. Based on the fossil records in and around the Tibetan Plateau, Equisetum could survive at a higher latitude during the Palaeogene and they could only grow in lower latitude since the Neogene, which might be the result of the tectonic deformation and climatic changes in the northern plateau since the Miocene. The current fossil, together with co-existed fossils, indicate that there have been rich drainages and temperate deciduous broad-leaved forest with grass in northwestern Qaidam Basin during the Oligocene.
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