Abstract During the last century, Grotta Romanelli (Southern Italy) has been a reference site for the European Late Pleistocene stratigraphy, due to its geomorphological setting and archaeological and palaeontological content. The beginning of the sedimentation inside the cave was attributed to MISs 5e and the oldest unearthed evidence of human occupation, including remains of hearths, was therefore referred to the Middle Palaeolithic. Recent surveys and excavations produced new U/Th dates, palaeoenvironmental interpretation and a litho-, morpho- and chrono-stratigraphical reassessment, placing the oldest human frequentation of the cave between MIS 9 and MIS 7, therefore embracing Glacial and Interglacial cycles. These new data provide evidence that the sea reached the cave during the Middle Pleistocene and human occupation occurred long before MISs 5e and persisted beyond the Pleistocene- Holocene boundary.
Suidae remains recovered from the late Pliocene site of Collepardo (Latium, central Italy) are described and assigned to Sus arvernensis, a small-sized Ruscinian to Early Villafranchian (MN14-MN16a) species. In Italy, S. arvernensis only occurs in the Triversa Faunal Unit (MN16a), supporting the recently revised chronology of Collepardo. CT-scan methods are used to virtually extract and analyse a newly discovered neurocranium, providing the content for the first inner cranial description carried out on an extinct Suidae. Our analysis reveals that S. arvernensis has an anteroposteriorly elongated and dorsoventrally flat cerebrum, similar to that of the Asian Babyrousa babyrussa and the African Hylochoerus meinertzhageni. These species substantially differ in size and are representatives of two widely diverging phylogenetic clades, excluding relatively simple evolutionary or allometric explanations for brain morphology in Suidae.
ABSTRACT In this paper, we present the results of the accelerator mass spectrometry radiocarbon (AMS 14 C) dating campaign performed on samples selected from different levels in Grotta Romanelli (Castro, Italy). Grotta Romanelli is one of the key sites for the chronology of Middle Pleistocene–Holocene in Mediterranean region. After the first excavation campaigns carried out in the first decades of the 1900s, the cave has been systematically re-excavated only since 2015. During the last excavation campaigns different faunal remains were selected and submitted for 14 C dating in order to confirm the chronology of the cave with a higher resolution. Isotopic ratio mass spectrometry (IRMS) measurements were also carried out on faunal remains.
Several craniodental and postcranial Bovinae fossil remains coming from the early Middle Pleistocene Contrada Monticelli (CM) site (Castellana Grotte, BA) and preserved at the Museum of Earth Sciences at the University of Bari are here studied in detail for the first time. In addition to Bovinae, the fauna of CM includes several Galerian taxa such as: Canis mosbachensis Soergel, 1925, Palaeoloxodon antiquus (Falconer & Cautley, 1847), Stephanorhinus hundsheimensis Toula, 1902, Equus sp. and Dama cf. roberti Breda & Lister, 2013. Although the scarcity of the sample and the lack of skulls, horncores or metapodial remains should suggest caution, the detailed morphological and biometric analyses allow to refer these remains to a large form of bison, here provisionally classified as Bison cf. schoetensacki Freudenberg, 1914. The presence of this species in the CM site would represent the southernmost occurrence of this taxon in Europe, validating the biochronological attribution of the CM site to the Isernia Faunal Unit (ca. 0.6 Ma). The presence of this bovid, and other large mammals in the site, provides further clues for the reconstruction of the paleoenvironment of Mediterranean Europe during the early Middle Pleistocene.
An updated description and revision of a left hemimandible assigned to Hyaenictitherium namaquensis, a dog-like hyaena from the late Miocene locality of As Sahabi (Libya, North Africa), is here provided. This fossil is part of the historical collection discovered by Carlo Petrocchi, the Italian researcher who excavated the site in the 1930s. The As Sahabi Hyaenictitherium represents the northernmost occurrence of H. namaquensis in Africa and one of the most informative evidence of this species known to date. The hemimandible has been digitised using photogrammetric techniques, morphological and biometric analyses have been carried out taking into account the African record of Hyaenictitherium.
Five rhinoceros species, mainly distinguishable through cranial, mandibular and dental morphology, were recognized during the Quaternary across Europe. The postcranial morphology of these taxa is quite variable, with strong superposition among species, especially between Stephanorhinus etruscus and Stephanorhinus hundsheimensis. Here, the complete sample including 25 postcranial bones from the early Middle Pleistocene site of Contrada Monticelli (Apulia, southern Italy) is described and compared through morphological and biometric analyses. A few bones, in addition to cranial and mandibular remains from this site, have been previously referred to Stephanorhinus hundsheimensis. We explored the morphological variability of Stephanorhinus hundsheimensis from Contrada Monticelli against the diagnostic characters described in the literature. The size variation of the Contrada Monticelli sample is also investigated, through a comparison with other Quaternary European rhinoceros samples. Our results indicate a wide morphological variability with several distinct characters and a strong morphological affinity between Stephanorhinus etruscus and S. hundsheimensis. Our results suggest that Stephanorhinus hundsheimensis from Contrada Monticelli is a species with a high degree of intraspecific variability, possibly a hybrid population or a population including two morphotypes of the same species.
The modern wolf, Canis lupus Linnaeus, 1758, has one of the largest ranges amongst carnivorans, and for this reason it shows local and regional differences for adaptation to a great variety of habitats, ranging from the arctic tundra to the Arabian desert. These differences are particularly evident as wolves follow the Bergmann's ecogeographical rule, with low latitude populations being generally smaller than those living at high latitudes. The fossil record of the modern wolf dates back to the second half of the Middle Pleistocene. The earliest records come from the French site Lunel-Viel 1 and are ascribed to the subspecies Canis lupus lunellensis Bonifay, 1971, and from the Italian Polledrara di Cecanibbio. These Middle Pleistocene forms were generally small in size, slightly larger than the last representative of Canis mosbachensis Soergel, 1925. During the last 400 kyr, European continental environments were profoundly affected by the glacial/interglacial cycles and much evidence suggests a trend toward a size increase for C. lupus. In particular, a number of these European large-sized wolves seem to be typical of cold phases, for example, Canis lupus maximus Boudadi-Maligne, 2012 from the OIS 3-2. The aim of the present study was to describe the morphological and morphometric variability of C. lupus from Avetrana bed 8 in comparison to other populations from northern and southern Italy, as well as from other localities in Europe, to obtain a better understanding of the biochronology, palaeobiogeography and evolution of this large carnivore in the last 125 kyr. The morphological and morphometric analyses pointed out the difference of C. lupus of Avetrana compared to the Apulian and to the Northern Italian wolves but also identified a marked affinity with the C. lupus of the glacial site Cardamone (Apulia), referred to OIS 2. This similarity support the idea of the dispersal of large morphotypes (glacial wolves) during cold phases.
(2022). Neurocranial anatomy of Sus arvernensis (Suidae, Mammalia) from Collepardo (Early Villafranchian; central Italy): taxonomic and biochronological implications. Historical Biology: Vol. 34, No. 1, pp. 108-120.
